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insect beetle macro photography nature wildlife biology entomology arthropods antennae legs wings exoskeleton thorax abdomen elytra mandibles maxillary palps compound eyes ocelli spiracle tarsi tibiae femora trochanters coxae prothorax mesothorax metathorax scutellum pygidium elytron membrane vein nervure cell spot pattern coloration variation adaptation camouflage mimicry defense predation pollination